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Hire Dr. Stuart S.
United Kingdom
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Scientific Editor & Writer | Emeritus Professor, Life & Health Sciences. Significant editorial & publishing experience.

Profile Summary
Subject Matter Expertise
Services
Writing Medical Writing, Non-Medical Regulatory Writing, Technical Writing, Business & Legal Writing, Copywriting, Creative Writing, General Proofreading & Editing, Translation
Work Experience

Professor, School of Life and Health Sciences

University of Roehampton

June 2013 - December 2022

Head of Research, School of Life and Health Sciences

University of Roehampton

January 2014 - September 2022

University of Roehampton

- August 2022

Education

Ph.D. Biological Sciences

University of Sussex

January 1995 - December 1998

Certifications
Publications
JOURNAL ARTICLE
Dominance style is a key predictor of vocal use and evolution across nonhuman primates @article{374fae7737e248a5868ff529822c7b7d, title = "Dominance style is a key predictor of vocal use and evolution across nonhuman primates", author = "Eithne Kavanagh and Street, {Sally E.} and Angwela, {Felix O.} and Bergman, {Thore J.} and Blaszczyk, {Maryjka B.} and Bolt, {Laura M.} and Margarita Brise{\~n}o-jaramillo and Michelle Brown and Chloe Chen-kraus and Zanna Clay and Camille Coye and Thompson, {Melissa Emery} and Alejandro Estrada and Claudia Fichtel and Barbara Fruth and Marco Gamba and Cristina Giacoma and Graham, {Kirsty E.} and Samantha Green and Grueter, {Cyril C.} and Shreejata Gupta and Gustison, {Morgan L.} and Lindsey Hagberg and Daniela Hedwig and Jack, {Katharine M.} and Kappeler, {Peter M.} and Gillian King-bailey and Barbora Kub{\v e}nov{\'a} and Alban Lemasson and Inglis, {David Macgregor} and Zarin Machanda and Andrew Macintosh and Bonaventura Majolo and Sophie Marshall and Stephanie Mercier and J{\'e}r{\^o}me Micheletta and Martin Muller and Hugh Notman and Karim Ouattara and Julia Ostner and Pavelka, {Mary S. M.} and Peckre, {Louise R.} and Megan Petersdorf and Fredy Quintero and Gabriel Ramos-fern{\'a}ndez and Robbins, {Martha M.} and Roberta Salmi and Isaac Schamberg and Schoof, {Val{\'e}rie A. M.} and Oliver Sch{\"u}lke and Stuart Semple and Silk, {Joan B.} and Sosa-lop{\'e}z, {J. Roberto} and Valeria Torti and Daria Valente and Raffaella Ventura and {Van De Waal}, Erica and Weyher, {Anna H.} and Claudia Wilke and Richard Wrangham and Christopher Young and Anna Zanoli and Klaus Zuberb{\"u}hler and Lameira, {Adriano R.} and Katie Slocombe", year = "2021", month = jul, day = "28", doi = "10.1098/rsos.210873", language = "English", volume = "8", journal = "Royal Society Open Science", issn = "2054-5703", publisher = "Royal Society, The", number = "7", } . Royal Society Open Science.
Intermittent pair-housing, pair relationship qualities, and HPA activity in adult female rhesus macaques @article{4a79f18ba1e94440876e5f5fe13a0db9, title = "Intermittent pair-housing, pair relationship qualities, and HPA activity in adult female rhesus macaques", abstract = "Laboratory rhesus macaques are often housed in pairs and may be temporarily or permanently separated for research, health, or management reasons. While both long‐term social separations and introductions can stimulate a stress response that impacts inflammation and immune function, the effects of short‐term overnight separations and whether qualities of the pair relationship mediate these effects are unknown. In this study, we investigated the effects of overnight separations on the urinary cortisol concentration of 20 differentially paired adult female rhesus macaques (Macaca mulatta) at the California National Primate Research Center. These females were initially kept in either continuous (no overnight separation) or intermittent (with overnight separation) pair‐housing and then switched to the alternate pair‐housing condition part way through the study. Each study subject was observed for 5 weeks, during which we collected measures of affiliative, aggressive, anxious, abnormal, and activity‐state behaviors in both pair‐housing conditions. Additionally, up to three urine samples were collected from each subject per week and assayed for urinary free cortisol and creatinine. Lastly, the behavioral observer scored each pair on four relationship quality attributes (“Anxious,” “Tense,” “Well‐meshed,” and “Friendly”) using a seven‐point scale. Data were analyzed using a generalized linear model with gamma distribution and an information theoretic approach to determine the best model set. An interaction between the intermittent pairing condition and tense pair adjective rating was in the top three models of the best model set. Dominance and rates of affiliation were also important for explaining urinary cortisol variation. Our results suggest that to prevent significant changes in HPA‐axis activation in rhesus macaque females, which could have unintended effects on research outcomes, pairs with “Tense” relationships and overnight separations preventing tactile contact should be avoided.{\textcopyright} 2018, Wiley Periodicals, Inc. The attached document (embargoed until 02/05/2019) is an author produced version of a paper published in AMERICAN JOURNAL OF PRIMATOLOGY uploaded in accordance with the publisher{\textquoteright}s self- archiving policy. The final published version (version of record) is available online at the link below. Some minor differences between this version and the final published version may remain. We suggest you refer to the final published version should you wish to cite from it.", author = "Darcy Hannibal and Lauren Cassidy and Jessica Vandeleest and Stuart Semple and Allison Barnard and Katie Chun and Sasha Winkler and Brenda McCowan", year = "2018", month = may, day = "2", doi = "10.1002/ajp.22762", language = "English", volume = "80", journal = "AMERICAN JOURNAL OF PRIMATOLOGY", issn = "0275-2565", publisher = "Wiley", number = "5", } . AMERICAN JOURNAL OF PRIMATOLOGY.
Primates’ behavioural responses to tourists @article{c2489be704714aa2a8a6f04b04a4d813, title = "Primates{\textquoteright} behavioural responses to tourists: evidence for a trade-off between potential risks and benefits", abstract = "The presence of, and interactions with tourists can be both risky and beneficial for wild animals. In wildlife tourism settings, animals often experience elevated rates of aggression from conspecifics, and they may also be threatened or physically aggressed by the tourists themselves. However, tourist provisioning of wild animals provides them with highly desirable foods. In situations of conflicting motivations such as this, animals would be expected to respond using behavioural coping mechanisms. In the present study, we investigated how animals respond to tourist pressure, using wild adult Barbary macaques in the Middle Atlas Mountains, Morocco, as a case study. We found evidence that these animals use a range of different behavioural coping mechanisms–physical avoidance, social support, affiliative, aggressive and displacement behaviours–to cope with the stress associated with tourists. The pattern of use of such behaviours appears to depend on a trade-off between perceived risks and potential benefits. We propose a framework to describe how animals respond to conflicting motivational situations, such as the presence of tourists, that present simultaneously risks and benefits.", author = "La{\"e}titia Mar{\'e}chal and Ann Maclarnon and Bonaventura Majolo and Stuart Semple", year = "2016", month = sep, day = "15", doi = "10.1038/srep32465", language = "English", volume = "6", journal = "Scientific Reports", issn = "2045-2322", publisher = "Nature Publishing Group", } . Scientific Reports.
Assessing the Effects of Tourist Provisioning on the Health of Wild Barbary Macaques in Morocco @article{98272c309b8f48dd83cfb48ee1540959, title = "Assessing the Effects of Tourist Provisioning on the Health of Wild Barbary Macaques in Morocco", abstract = "Feeding wildlife is a very popular tourist activity, largely because it facilitates the close observation of animals in their natural habitat. Such provisioning may benefit animals by improving their survival and reproductive success, especially during periods of natural food shortage. However, provisioning by tourists may also have negative impacts on the health of the animals involved; to date such impacts are poorly understood. Here, we investigated the effects of tourist provisioning on the health of wild adult Barbary macaques, Macaca sylvanus, in Morocco. We compared health measures between a heavily provisioned group and a group that received negligible food from tourists and, in the former group, we also assessed health measures in relation to the intensity of provisioning. We used a broad range of non-invasive health measures relating to birth rate and survival, disease and injury risk, body size and condition, and physiological stress. Our findings indicate that feeding by tourists may overall have negative impacts on the health of Barbary macaques, being linked in particular to larger body size, elevated stress levels and more alopecia. Finally, we propose a framework to help consider the potential costs and benefits of provisioning, which may facilitate future research and management decisions on whether—and how much—provisioning is acceptable.", author = "Latitia Marechal and Stuart Semple and Bonaventura Majolo and Ann MacLarnon", note = "{\textcopyright} 2016 Mar{\'e}chal et al. This is the final published version of the article (version of record) uploaded in accordance with the publisher{\textquoteright}s self-archiving policy. It first appeared online at http://dx.doi.org/10.1371/journal.pone.0155920. ", year = "2016", month = may, day = "20", doi = "10.1371/journal.pone.0155920", language = "English", volume = "11", journal = "PLoS One", issn = "1932-6203", publisher = "Public Library of Science", number = "5", } . PLoS One.
Gelada vocal sequences follow Menzerath's linguistic law @article{8630db1072c44050bc31ad47d1dc4000, title = "Gelada vocal sequences follow Menzerath's linguistic law", abstract = "Identifying universal principles underpinning diverse natural systems is a key goal of the life sciences. A powerful approach in addressing this goal has been to test whether patterns consistent with linguistic laws are found in nonhuman animals. Menzerath's law is a linguistic law that states that, the larger the construct, the smaller the size of its constituents. Here, to our knowledge, we present the first evidence that Menzerath's law holds in the vocal communication of a nonhuman species. We show that, in vocal sequences of wild male geladas (Theropithecus gelada), construct size (sequence size in number of calls) is negatively correlated with constituent size (duration of calls). Call duration does not vary significantly with position in the sequence, but call sequence composition does change with sequence size and most call types are abbreviated in larger sequences. We also find that intercall intervals follow the same relationship with sequence size as do calls. Finally, we provide formal mathematical support for the idea that Menzerath's law reflects compression-the principle of minimizing the expected length of a code. Our findings suggest that a common principle underpins human and gelada vocal communication, highlighting the value of exploring the applicability of linguistic laws in vocal systems outside the realm of language.", author = "Gustison, {Morgan L} and Stuart Semple and Ramon Ferrer-I-Cancho and Bergman, {Thore J}", year = "2016", month = may, day = "10", doi = "10.1073/pnas.1522072113", language = "English", volume = "113", pages = "E2750--E2758", journal = "PNAS: Proceedings of the National Academy of Sciences", issn = "0027-8424", publisher = "National Acad Sciences", number = "19", } . PNAS: Proceedings of the National Academy of Sciences.
Stuart Semple, Kevin A. Rosenfield, Constance Dubuc, Alexander V. Georgiev, Dario Maestripieri(2016). Evidence that male face shape in rhesus macaques (Macaca mulatta) is subject to intersexual selection . AMERICAN JOURNAL OF PHYSICAL ANTHROPOLOGY. 159. p. 273--273. Wiley
Stuart Semple, Amanda M. Maijer(2016). Investigating Potential Effects of the Contraceptive Implanon on the Behavior of Free-Ranging Adult Female Barbary Macaques . JOURNAL OF APPLIED ANIMAL WELFARE SCIENCE. 19. (1). p. 16--23. Routledge: Taylor & Francis Group
Stuart Semple, Lauren Cassidy, Darcy Hannibal, Brenda McCowan(2015). Behavioural and Physiological Effects of Housing Type on Laboratory Housed Female Rhesus Macaques (Macaca mulatta) . Folia Primatologica. 86. (4). p. 259--260. Karger
Stuart Semple, Charlotte Carne, Julia Lehmann(2015). Investigating constraints on the survival of orangutans across Borneo and Sumatra . TROPICAL CONSERVATION SCIENCE. 8. (4). p. 940--954. Mongabay.com
Stuart Semple, Changiz Mohiyeddini, Stephanie Bauer(2015). Neuroticism and stress: . Anxiety Stress and Coping. 28. (4). p. 391--407. Routledge: Taylor & Francis Group
Stuart Semple, Kevin Rosenfield, Constance Dubuc(2015). Testing for Sexual Selection on Rhesus Macaque (Macaca mulatta) Face Shape . Folia Primatologica. 86. (4). p. 349--349. Karger
Stuart Semple, S. C. Faulkner, M. D. Stevenson, R. Verity, A. H. Mustari, D. G. Tosh, S. C. Le Comber(2015). Using geographic profiling to locate elusive nocturnal animals: . JOURNAL OF ZOOLOGY. 295. (4). p. 261--268. Wiley-Blackwell
Stuart Semple, Ramon Ferrer-I-Cancho, Thore Bergman, Minna Hsu, Govindasamy Agoramoorthy, Morgan Gustison(2015). Linguistic Laws in Primate Vocal Communication . Folia Primatologica. 86. (4). p. 357--357. Karger
Stuart Semple, Charlotte Carne, Helen Morrogh-Bernard, Klaus Zuberbuehler, Julia Lehmann(2014). The Risk of Disease to Great Apes: . PLoS One. 9. (4). Public Library of Science
Stuart Semple, Lauren J. N. Brent, Ann MacLarnon, Angelina Ruiz-Lambides, Janis Gonzalez-Martinez, Michael L. Platt(2014). Personality Traits in Rhesus Macaques (Macaca mulatta) Are Heritable but Do Not Predict Reproductive Output . INTERNATIONAL JOURNAL OF PRIMATOLOGY. 35. (1). p. 188--209. Springer
Stuart Semple, Chiara Borg, Bonaventura Majolo, Mohamed Qarro(2014). A Comparison of Body Size, Coat Condition and Endoparasite Diversity of Wild Barbary Macaques Exposed to Different Levels of Tourism . Anthrozoos. 27. (1). p. 49--63. Routledge: Taylor & Francis Group
Stuart Semple, Emma K. Wallace, Mark Kingston-Jones, Matthew Ford(2013). An investigation into the use of music as potential auditory enrichment for moloch gibbons (Hylobates moloch) . Zoo Biology. 32. (4). p. 423--426. Wiley
Stuart Semple, Ramon Ferrer-i-Cancho, Antoni Hernandez-Fernandez, David Lusseau, Govindasamy Agoramoorthy, Minna J. Hsu(2013). Compression as a Universal Principle of Animal Behavior . COGNITIVE SCIENCE. 37. (8). p. 1565--1578. Wiley
Stuart Semple, Changiz Mohiyeddini, Stephanie Bauer(2013). Displacement Behaviour Is Associated with Reduced Stress Levels among Men but Not Women . PLoS One. 8. (2). Public Library of Science
Stuart Semple, Changiz Mohiyeddini(2013). Displacement behaviour regulates the experience of stress in men . STRESS-THE INTERNATIONAL JOURNAL ON THE BIOLOGY OF STRESS. 16. (2). p. 163--171. Routledge: Taylor & Francis Group
Stuart Semple, Christy Harrison, Julia Lehmann(2013). Grooming and Anxiety in Barbary Macaques . ETHOLOGY. 119. (9). p. 779--785. Wiley
Stuart Semple, Richard McFarland, Ann MacLarnon, Michael Heistermann(2013). Physiological stress hormone levels and mating behaviour are negatively correlated in male rhesus macaques (Macaca mulatta) . Animal Biology. 63. (3). p. 331--341. Brill Academic Publishers
Stuart Semple, Charlotte Carne, Helen Morrogh-Bernard, Klaus Zuberbuehler, Julia Lehmann(2013). Predicting the Vulnerability of Great Apes to Disease: . PLoS One. 8. (12). Public Library of Science
Stuart Semple, Changiz Mohiyeddini, Stephanie Bauer(2013). Public self-consciousness moderates the link between displacement behaviour and experience of stress in women . STRESS-THE INTERNATIONAL JOURNAL ON THE BIOLOGY OF STRESS. 16. (4). p. 384--392. Routledge: Taylor & Francis Group
Stuart Semple, Lauren J. N. Brent, Ann MacLarnon, Michael L. Platt(2013). Seasonal changes in the structure of rhesus macaque social networks . Behavioral Ecology and Sociobiology. 67. (3). p. 349--359. Springer
Stuart Semple, Morgan L. Gustison, Ann MacLarnon, Sue Wiper(2012). An experimental study of behavioural coping strategies in free-ranging female Barbary macaques (Macaca sylvanus) . STRESS-THE INTERNATIONAL JOURNAL ON THE BIOLOGY OF STRESS. 15. (6). p. 608--617. Routledge: Taylor & Francis Group
Stuart Semple, E. J. Bethell, Amanda Holmes, Ann MacLarnon(2012). Cognitive bias in a non-human primate: . Animal Welfare. 21. (2). p. 185--195. Universities Federation for Animal Welfare
Stuart Semple, Stefano S. K. Kaburu, Ann MacLarnon, Bonaventura Majolo, Mohamed Qarro(2012). Dominance rank and self-scratching among wild female Barbary macaques (Macaca sylvanus) . African Zoology. 47. (1). p. 74--79. Routledge: Taylor & Francis Group
Stuart Semple, Emily J. Bethell, Amanda Holmes, Ann MacLarnon(2012). Evidence That emotion mediates social attention in Rhesus Macaques . PLoS One. 7. (8). Public Library of Science
Stuart Semple, Victoria Buckley(2012). Evidence that displacement activities facilitate behavioural transitions in ring-tailed lemurs . BEHAVIOURAL PROCESSES. 90. (3). p. 433--435. Elsevier
Stuart Semple, Odile Naim, C Tourrand, L Bigot, C Cadet, B Cauvin, L. Montaggioni, H Bruggemann(2012). Fringing reefs of Reunion Island. Part 1: . Atoll Research Bulletin. Smithsonian Institution
Stuart Semple, Julia Fischer, Gisela Fickenscher, Rebecca Juergens, Eberhard Kruse, Michael Heistermann, Ofer Amir(2011). Do Women's Voices Provide Cues of the Likelihood of Ovulation? . PLoS One. 6. (9). Public Library of Science
Stuart Semple, Laetitia Marechal, Bonaventura Majolo, Mohamed Qarro, Michael Heistermann, Ann MacLarnon(2011). Impacts of tourism on anxiety and physiological stress levels in wild male Barbary macaques . BIOLOGICAL CONSERVATION. 144. (9). p. 2188--2193. Elsevier
Stuart Semple, J. R. Rogge(2011). SEX DIFFERENCES IN SOCIALITY AND SOCIAL BOND MAINTENANCE AMONG WILD OLIVE BABOONS . AMERICAN JOURNAL OF PRIMATOLOGY. 73. p. 109--109. Wiley
Stuart Semple, L. J. N. Brent, C. Dubuc, Michael Heistermann, Ann MacLarnon(2011). Social capital and physiological stress levels in free-ranging adult female rhesus macaques . PHYSIOLOGY & BEHAVIOR. 102. (1). p. 76--83. Elsevier
Stuart Semple, Jacklyn J. Ellis, Ann MacLarnon, Michael Heistermann(2011). The social correlates of self-directed behaviour and faecal glucocorticoid levels among adult male olive baboons (Papio hamadryas anubis) in Gashaka-Gumti National Park, Nigeria . African Zoology. 46. (2). p. 302--308. Routledge: Taylor & Francis Group
Stuart Semple, Kara L. Moses(2011). Primary seed dispersal by the black-and-white ruffed lemur (Varecia variegata) in the Manombo forest, south-east Madagascar . JOURNAL OF TROPICAL ECOLOGY. 27. p. 529--538. Cambridge University Press
Stuart Semple, Minna J. Hsu, Govindasamy Agoramoorthy(2010). Efficiency of coding in macaque vocal communication . BIOLOGY LETTERS. 6. (4). p. 469--471. Royal Society, The
Stuart Semple, Lisa Reamer, Zena Tooze, Claire Coulson(2010). Correlates of self-directed and stereotypic behaviours in captive red-capped mangabeys (Cercocebus torquatus torquatus) . Applied Animal Behaviour Science. 124. (42401). p. 68--74. Elsevier
Stuart Semple, Tabitha Price, Kate Arnold, Klaus Zuberbuehler(2009). Pyow but not hack calls of the male putty-nosed monkey (Cercopithcus nictitans) convey information about caller identity . BEHAVIOUR. 146. p. 871--888. Brill Academic Publishers
Stuart Semple, James P. Higham, Ann MacLarnon, Michael Heistermann, Caroline Ross(2009). Rates of self-directed behaviour and faecal glucocorticoid levels are not correlated in female wild olive baboons (Papio hamadryas anubis) . STRESS-THE INTERNATIONAL JOURNAL ON THE BIOLOGY OF STRESS. 12. (6). p. 526--532. Routledge: Taylor & Francis Group
Stuart Semple, Nathalie C. Greeno(2009). Sex differences in vocal communication among adult rhesus macaques . EVOLUTION AND HUMAN BEHAVIOR. 30. (2). p. 141--145. Elsevier
Stuart Semple, Constance Dubuc, Lauren J. N. Brent, Amanda K. Accamando, Melissa S. Gerald, Ann MacLarnon, Michael Heistermann, Antje Engelhardt(2009). Sexual skin color contains information about the timing of the fertile phase in free-ranging Macaca Mulatta . INTERNATIONAL JOURNAL OF PRIMATOLOGY. 30. (6). p. 777--789. Springer
Stuart Semple, James P. Higham, Ann MacLarnon, Michael Heistermann, Caroline Ross(2009). Female reproductive signaling, and male mating behavior, in the olive baboon . HORMONES AND BEHAVIOR. 55. (1). p. 60--67. Elsevier
Stuart Semple, Jane Ebert Haakonsson(2009). Lateralisation of trunk movements in captive Asian elephants (Elephas maximus) . LATERALITY. 14. (4). p. 413--422. Routledge: Taylor & Francis Group
Stuart Semple, James P. Higham, Ann MacLarnon, Caroline Ross, Michael Heistermann(2008). Baboon sexual swellings: . HORMONES AND BEHAVIOR. 53. (3). p. 452--462. Elsevier
Stuart Semple, Ann MacLarnon, K. Shutt, M Heistermann, J. P. Higham(2008). Sociality and stress in Macaques and baboons . Folia Primatologica. 79. (5). p. 355--355. Karger
Stuart Semple, James P. Higham, Michael Heistermann, Caroline Ross, Ann MacLarnon(2008). The timing of ovulation with respect to sexual swelling detumescence in wild olive baboons . PRIMATES. 49. (4). p. 295--299. Springer
Stuart Semple, Gemma Carder(2008). Visitor effects on anxiety in two captive groups of western lowland gorillas . Applied Animal Behaviour Science. 115. (42463). p. 211--220. Elsevier
Stuart Semple, Susan M. Wiper(2007). The function of teeth chattering in male Barbary Macaques (Macaca sylvanus) . AMERICAN JOURNAL OF PRIMATOLOGY. 69. (10). p. 1179--1188. Wiley
Stuart Semple, Kathryn Shutt, Ann MacLarnon, Michael Heistermann(2007). Grooming in Barbary macaques: . BIOLOGY LETTERS. 3. (3). p. 231--233. Royal Society, The
Stuart Semple, Anna H. Weyher, Caroline Ross(2006). Gastrointestinal parasites in crop raiding and wild foraging Papio anubis in Nigeria . INTERNATIONAL JOURNAL OF PRIMATOLOGY. 27. (6). p. 1519--1534. Springer
Stuart Semple, Chris Sandbrook(2006). The rules and the reality of mountain gorilla Gorilla beringei beringei tracking: . ORYX. 40. (4). p. 428--433. Cambridge University Press
Stuart Semple, K McComb(2005). Coevolution of vocal communication and sociality in primates . BIOLOGY LETTERS. 1. (4). p. 381--385. Royal Society, The
Stuart Semple(2002). Immune system evolution among anthropoid primates: . Proceedings of The Royal Society B Biological Sciences. Royal Society, The
Stuart Semple(2002). Information content of female copulation calls in yellow baboons . AMERICAN JOURNAL OF PRIMATOLOGY. Wiley
OTHER
Stuart Semple, Nienke Alberts, Julia Lehmann(2015). Quantifying Fission-Fusion Dynamics: . Folia Primatologica. 86. (4). p. 236--236. Karger
Stuart Semple, C. Harrison, S. Wiper, Julia Lehmann(2011). Grooming and Anxiety in Female Barbary Macaques . Folia Primatologica. 82. (6). p. 325--326. Karger
BOOK CHAPTER
Stuart Semple, Bonaventura Majolo, Els van Lavieren, Laëtitia Maréchal, Ann MacLarnon, Garry Marvin, Mohamed Qarro(2013). Out of Asia: . The Macaque Connection. p. 167--183. Springer
Stuart Semple, Nienke Alberts, Julia Lehmann(2013). The effect of filtering on animal networks . "Social Networking:. Nova
JOURNAL ISSUE
Stuart Semple, J. Higham(2013). Primate Signals . AMERICAN JOURNAL OF PRIMATOLOGY. 75. p. 613--620. Wiley